Heliobolus lugubris   (SMITH, 1838)

Heliobolus lugubris   (SMITH, 1838)


          
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  • Alexander, G. & Marais, J. (2007) -  A Guide to the Reptiles of Southern Africa -  Struik publishers, Cape Town.  

  • Arnold, E.N. (2002) -  History and function of scale microornamentation in lacertid lizards. -  Journal of Morphology, 252 (2): 145-169.    

  • Auerbach, R.D. (1986) -  First steps in Setswana herpetology. -  Botswana Notes and Records Volume, 18: 71-90.  

  • Baeckens, S. & Martin, S. & Garcia-Roa, R. & Pafilis, P. & Huyghe, K. & Damme, R. van (2017) -  Environmental conditions shape the chemical signal design of lizards. -  Functional Ecology, 32 (2): 566-580.    

  • Baptista, N. & António, T. & Branch, W.R. (2019) -  The herpetofauna of Bicuar National Park and surroundings, southwestern Angola: a preliminary checklist. -  Amphibian & Reptile Conservation, 13 (2): 96-130.    

  • Barts, M. & W.D. Haacke (1997) -  Zur Reptilienfauna der Tsodilo-Berge und angrenzender Gebiete in NW-Botswana – Teil I: Testudines: Testudinidae; Sauria I. -  Sauria, Berlin, 19 (1): 3-12.  

  • Bates, M.F. (1991) -  New distribution records for amphibians and reptiles from the Cape Province, South Africa. -  Journal of the Herpetological Association of Africa, Johannesburg, South Africa, 39 : 14-18.    

  • Bates, M.F. & Branch, W.R. & Bauer, A.M. & Burger, M. & Marais, J. & Alexander, G.J. & Villiers, M.S. de (eds.) (2014) -  Atlas and Red List of the Reptiles of South Africa, Lesotho and Swaziland. -  Suricata 1. South African National Biodiversity Institute. Pretoria. 485 pp.  

  • Bauer, A.M. & Kennedy, A.M. & Lewis, P.J. & Thies, M.L. & Gabadirwe, M. (2009) -  The herpetofauna of Koanaka South and adjacent regions, Ngamiland, Botswana. -  Herpetological Bulletin, 107: 16-26.    

  • Bennett, A.F. & Huey, R.B. & John-Alder, H. (1984) -  Physiological correlates of natural activity and locomotor capacity in two species of lacertid lizards. -  Journal of Comparative Physiology B, Berlin, 154 (2): 113-118.     

  • Bocage, J.V.B. du (1867) -  Segunda lista dos reptis das possessões portuguezas d’Africa occidental que existem no Museu de Lisboa. -  Jornal de sciencias mathematicas, physicas e naturaes, Academia Real das Sciencias de Lisboa, I (3): 217-228.  

  • Bocage, J.V.B. du (1895) -  Herpétologie d’Angola et du Congo. -  Ministério da Marinha e das Colónias, Lisbonne. 203 pp.  

  • Boettger, O. (1893) -  Katalog der Reptilien-Sammlung im Museum der Senckenbergischen Naturforschenden Gesellschaft in Frankfurt am Main, I. Teil (Phynchocephalen, Schildkröten, Krokodile, Eidechsen, Chamäleons). -  Gebrüder Knauer, Frankfurt. 140 pp.  

  • Boulenger, G.A. (1905) -  A List of the Batrachians and Reptiles collected by Dr. W.J. Ansorge in Angola, with descriptions of new species. -  The Annals and magazine of natural history, (7) 16: 105-115.  

  • Boulenger, G.A. (1906) -  Revised List of South African Reptiles and Batrachians. Family Lacertidae. -  Annals of the South African Museum, Cape Town, South Africa, 5: 473-479.  

  • Branch, B. (1988) -  Field guide to snakes and other reptiles of southern Africa. -  Struik Publishers, Cape Town. 328 pp.  

  • Branch, W.R. (1981) -  An annotated checklist of the lizards of the Cape Province, South Africa. -  Annals of the Cape Provincial Museums Natural History, 13 (11): 141-165.    

  • Branch, W.R. & Baard, E.H.W. & Haacke, W.D. & Jacobsen, N. & Ponton, J.C. & Broadley, D.G. (1988) -  A provisional and annotated checklist of the herpetofauna of southern africa. -  Journal of the Herpetological Association of Africa, Johannesburg, South Africa, 34 (1): 1-19.  

  • Branch, W.R. & Vaz Pinto, P. & Baptista, N. & Conradie, W. (2019) -  The Reptiles of Angola: History, Diversity, Endemism and Hotspots. -  In: Huntley B., Russo V., Lages F., Ferrand N. (eds.) Biodiversity of Angola: 283-334.    

  • Broadley, D.G. (1961) -  A provisional check list of the crocodiles, chelonians and lizards of Rhodesia and Nyasaland. -  The Journal of the Herpetological Association of Rhodesia, Umtali, Zimbabwe, 16: 3-8.  

  • Broadley, D.G. (1965) -  A provisional checklist of the lizards of Rhodesia. -  The Journal of the Herpetological Association of Rhodesia, Umtali, Zimbabwe, 23/24: 7-12.  

  • Broadley, D.G. (1965) -  A provisional key to the lizards of Rhodesia. -  The Journal of the Herpetological Association of Rhodesia, Umtali, Zimbabwe, 23/24: 12-19.  

  • Broadley, D.G. (1967) -  An Expedition to the South-Western Kalahari, January-March, 1967. -  Journal of the Herpetological Association of Africa, Johannesburg, South Africa, 3 (1): 19-30.  

  • Broadley, D.G. (1967) -  An Umtali Museum Expedition to the Gona-Re-Zhou, December, 1966. -  Journal of the Herpetological Association of Africa, Johannesburg, South Africa, 3 (1): 14-18.  

  • Broadley, D.G. (1978) -  English names for Rhodesian reptiles and amphibians. -  Journal of the Herpetological Association of Africa, Johannesburg, South Africa, 19 (1): 25-36.  

  • Broadley, D.G. (2004) -  Herpetofauna of the four corners area. -  In: Timberlake, J.R. & Childes, S.L. (Eds.) 2004. Biodiversity of the Four Corners Area: Technical Reviews Volume Two (Chapters 5-15). Occasional Publications in Biodiversity No 15, Biodiversity Foundation for Africa, Bulawayo/Zambezi Society, Harare, Zimbabwe: 313- 346.    

  • Brown, T.K. & Nagy, K.A. (2007) -  Lizard energetics and the sit-and-wait vs. wide-foraging paradigm. -  In: Lizard ecology: the evolutionary consequences of foraging mode. Cambridge University Press: 120-140.  

  • Castilla, A.M. & Van Damme, R. & Bauwens, D. (1999) -  Field body temperatures, mechanisms of thermoregulation and evolution of thermal characteristics in lacertid lizards. -  Natura Croatica, Zagreb, 8 (3): 253-274.    

  • Ceríaco, L.M.P. & Sá, D.d.A.C. de & Bandeira, S. & Valério, H. & Stanley, E.L. & Kuhn, A.L. & Marques, M.P. & Vindum, J.V. & Blackburn, D.C. & Bauer, A.M. (2016) -  Herpetological Survey of Iona National Park and Namibe Regional Natural Park, with a Synoptic List of the Amphibians and Reptiles of Namibe Province, Southwestern Angola. -  Proceedings of the California Academy of Sciences, series 4, 63 (2): 15-61.    

  • Conradie, W. & Bills, R. & Branch, W.R. (2016) -  The herpetofauna of the Cubango, Cuito, and lower Cuando river catchments of south-eastern Angola. -  Amphibian and Reptile Conservation, 10 (2): 6-36.    

  • Conradie, W. & Engelbrecht, H.M. & Herrel, A. & Measey, G.J. & Nielsen, S.V. & Vanhooydonck, B. & Tolley, K.A. (2011) -  Reptile survey of Venetia Limpopo Nature Reserve, Limpopo province - South Africa. -  African Herp News, Bloemfontein, South Africa, 55: 6-12.  

  • Cunningham, P.L. (2011) -  Vertebrate Fauna and Flora associated with the Desert Star South–Phases 1A-C–Area -  Environment and Wildlife Consulting Namibia  

  • Dubke, M. & Hipsley, C.A. & Müller, J. (2018) -  Comparative skull osteology and preliminary systematic revision of the African lizard genus Heliobolus (Squamata: Lacertidae). -  African Journal of Herpetology, Johannesburg, South Africa, 67 (2): 160-197.    

  • Duméril, A.M.C. & Bibron, G. (1839) -  Livre quatrième: De l`ordre des lézards ou des sauriens - Seconde sous-famille. Autosaures Coelodontes. -  Erpétologie Générale ou Histoire Naturelle Complète des Reptiles, 5: 153-317.  

  • Engleder, A. & Haring, E. & Kirchhof, S. & Mayer, W. (2013) -  Multiple nuclear and mitochondrial DNA sequences provide new insights into the phylogeny of South African Lacertids (Lacertidae, Eremiadinae). -  Journal of Zoological Systematics and Evolutionary Research, Berlin, 51 (2): 132-143.    

  • Fitzinger, L.I. (1843) -  II. Sectio. Coelodontes. Dum. Bibr. -  In: Systema Reptilium, Fasciculus Primus: Amblyglossae. Braumüller & Seidl, Wien: 20-21.  

  • FitzSimons, V. (1935) -  Scientific results of the Vernay-Lang Kalahari Expedition, March to September, 1930. Reptilia and Amphibia. Part 2. -  Annals of the Transvaal Museum, 16 (2): 295-397.  

  • FitzSimons, V. (1937) -  Notes on the reptiles and amphibians collected and described from South Africa by Andrew Smith. -  Annals of the Transvaal Museum, 17 (4): 259-274.    

  • Fitzsimons, V.F.M. (1938) -  Transvaal Museum Expedition to South-West Africa and Little Namaqualand, May to August 1937 - Reptiles and Amphibians. -  Annals of the Transvaal Museum, 19 (2): 153-209.   

  • Fuchs, J. & Grohmann, C. & Rödel, M.-O. & Penner, J. (2006) -  Ökologische Untersuchungen an Reptilien einer Dornbuschsavanne in Zentralnamibia – Unwägbarkeiten der Freilandforschung. -  elaphe, 14 (4): 45-50.  

  • Goldberg, S.R. (2006) -  Fortpflanzungszyklus der Bushveld-Eidechse Heliobolus lugubris (Squamata: Lacertidae) im südlichen Afrika. -  Der Salamander, Rheinbach, 2 (2/3): 151-154.  

  • Goldberg, S.R. (2006) -  Reproductive cycle of the bushveld lizard Heliobolus lugubris (Squamata: Lacertidae) from southern Africa. -  Salamandra, 42 (2/3): 151-154.    

  • Gow, C.E. (1962) -  Some collecting techniques and general notes. -  The Journal of the Herpetological Association of Rhodesia, Umtali, Zimbabwe, 19: 13-14.  

  • Griffin, M. (2002) -  Annotated Checklist and Provisional Conservation Status of Namibian Reptiles. -  Technical Reports of Scientific Services Nr. 1. Windhoek: Ministry of Environment and Tourism. 76 pp.    

  • Haacke, W.D. (1970) -  New herpetological records from South West Africa. -  Annals of the Transvaal Museum, 26: 277-283.  

  • Haacke, W.D. (1984) -  The herpetology of the southern Kalahari domain. -  Koedoe - African Protected Area Conservation and Science, 27 (1): 171-186.    

  • Herrmann, H.-W. & Branch, W.R. (2013) -  Fifty years of herpetological research in the Namib Desert and Namibia with an updated and annotated species checklist. -  Journal of Arid Environments, 93: 94-115.    

  • Hewitt, J. (1910) -  A key to the South African species of Geckonidae, Scincidae, Gerrhosauridae, and Lacertidae, together with some notes on the specific characters and a brief summary of the known of their distribution. -  Annals of the Transvaal Museum, 2: 77-115.   

  • Hipsley, C. (2012) -  Evolutionary consequences of Cenozoic climate change on African lacertid lizards (Squamata: Lacertidae). -  Ph.D. dissertation, University of California. 126 pp.    

  • Hoesch, W. (1961) -  Südwestafrikanische Echsen in Freiheit und in Gefangenschaft. -  Die Aquarien- und Terrarien-Zeitschrift, 14 (9): 278-280; 14 (10): 307-309.   

  • Huey, R.B. & Pianka, E.R. (1977) -  Natural selection for juvenile lizards mimicking beetles. -  Science, 195: 201-203.  

  • Huey, R.B. & Pianka, E.R. (1977) -  Seasonal Variation in Thermoregulatory Behavior and Body Temperature of Diurnal Kalahari Lizards. -  Ecology, 58 (5): 1066-1075.     

  • Huey, R.B. & Pianka, E.R. (1981) -  Ecological Consequences of Foraging Mode. -  Ecology, 62 (4 ): 991-999.    

  • Huey, R.B. & Pianka, E.R. (2007) -  Historical introduction: on widely foraging for Kalahari lizards. -  In: Lizard ecology: the evolutionary consequences of foraging mode. Cambridge University Press.  

  • Huey, R.B. & Pianka, E.R. (2018) -  Body temperature distributions of active diurnal lizards in three deserts: Skewed up or skewed down? -  Functional Ecology, 32 (2): 334-344.    

  • Huey, R.B. & Pianka, E.R. & Hoffmann, J.A. (1977) -  Seasonal variation in thermoregulatory behavior and body temperature of diurnal Kalahari lizards. -  Ecology, 58: 1066-1075.     

  • Huey, R.B. & Pianka, E.R. & Vitt, L.J. (2001) -  How often do lizards “Run on empty“? -  Ecology, 82 (1): 1-7.    

  • Jacobsen, N. (1978) -  Colloquial names for southern African reptiles and amphibians. -  Journal of the Herpetological Association of Africa, Johannesburg, South Africa, 18 (1): 7-13.  

  • Kennedy, A.M. & Marais, J. & Bauer , A.M. & Lewis, P.J. & Thies, M.L. (2012) -  Effect of Fire on the Herpetofauna of the Koanaka Hills, Ngamiland, Botswana. -  Check List, 8 (4): 666-674.    

  • Kirchhof, S. (2011) -  Auf der Suche nach Lacertiden in Südafrika: Ergebnisse einer Reise in den Nordosten des Landes.  -  Die Eidechse, Magdeburg/Hamburg, 22 (3): 71-82.   

  • Kirchhof, S. & Krämer, M. & Linden, J. & Richter, K. (2010) -  The reptile species assemblage of the Soutpansberg (Limpopo Province, South Africa) and its characteristics. -  Salamandra, 46 (3): 147-166.    

  • Kirchhof, S. & Rödel, M.-O. & Mueller, J. (2016) -  Does climate change drive extinction risk in Namibian lizards (Lacertidae). -  Oral presentation, 12th Conference Herpetological Association of Africa. – African Herp News, Bloemfontein, South Africa, 63: 38.    

  • Lewin, A. & Feldman, A. & Bauer, A.M. & Belmaker, J. & Broadley, D.G. & Chirio, L. & Itescu, Y. & LeBreton, M. & Maza, E. & Meirte, D. & Nagy, Z.T. & Novosolov, M. & Roll, U. & Tallowin, O. & Trape, J.-F. & Vidan, E. & Meiri, S. (2016) -  Patterns of species richness, endemism and environmental gradients of African reptiles. -  Journal of Biogeography, 43 (12): 2380-2390.    

  • Marais, J. (2009) -  Trip to Koanaka Hills, Botswana. -  http://www.johanmarais.co.za    

  • Marques, M.P. & Ceríaco, L.M.P. & Blackburn, D.C. & Bauer, A.M. (2018) -  Diversity and distribution of the amphibians and terrestrial reptiles of Angola: Atlas of historical and bibliographic records (1840-2017). -  Proceedings of the California Academy of Sciences, Series 4, 65, Supplement II: 1-501.    

  • Marques, M.P. & Ceríaco, L.M.P. & Heinicke, M.P. & Chehouri, R.M. & Conradie, W. & Tolley, K.A. & Bauer, A.M. (2022) -  The Angolan bushveld lizards, genus Heliobolus Fitzinger, 1843 (Squamata: Lacertidae): Integrative taxonomy and the description of two new species. -  Vertebrate Zoology, 72: 745–769 (DOI 10.3897/vz.72.e85269).    

  • Mayer, W. & Benyr, G. (1994) -  Albumin-Evolution und Phylogenese in der Familie Lacertidae. -  Annalen des Naturhistorischen Museums in Wien, 96 B: 621-648.    

  • Mayer, W. & Berger-Dell´Mour, H. (1988) -  Proteinelektrophoretische Untersuchungen zur Systematik der Gattungen Aporosaura, Meroles, Pedioplanis und Heliobolus (Sauria: Lacertidae) aus Südwest-Afrika. -  Herpetozoa, Wien, 1 (1/2): 23-29.    

  • Mayer, W. & Pavlicev, M. (2007) -  The phylogeny of the family Lacertidae (Reptilia) based on nuclear DNA sequences: Convergent adaptations to arid habitats within the subfamily Eremiainae. -  Molecular Phylogenetics and Evolution, 44: 1155–1163.    

  • Mayer, W. & Richter, K. (1990) -  Die Wüstenrenner-Eidechsen Namibias – Vorkommen, Pflege und Zucht. -  herpetofauna, 12 (66): 21-30.  

  • McAllister, C.T. & Bursey, C.R. & Freed, P.S. (2011) -  Endoparasites (Cestoidea, Nematoda, Pentastomida) of Reptiles (Sauria, Ophidia) from the Republic of Namibia. -  Comparative Parasitology, Washington, 78 (1): 140-151.    

  • McBrayer, L.D. (2004) -  The relationship between skull morphology, biting performance and foraging mode in Kalahari lacertid lizards. -  Zoological Journal of the Linnean Society, 140 (3): 403-416.    

  • McBrayer, L.D. & Wylie, J.E. (2009) -  Concordance between locomotor morphology and foraging mode in lacertid lizards -  Zoology, 112 (5): 370–378.    

  • Meiri, S. (2008) -  Evolution and ecology of lizard body sizes. -  Global Ecology and Biogeography, 17 (6): 724-734.    

  • Meiri, S. & Bauer, A.M. & Chirio, L. & Colli, G.R. & Das, I. & Doan, T.M. & Feldman, A. & Herrera, F.-C. & Novosolov, M. & Pafilis, P. Pincheira-Donoso, D. & Powney, G. & Torres-Carvajal, O. & Uetz, P. & Van Damme, R. (2013) -  Are lizards feeling the heat? A tale of ecology and evolution under two temperatures. -  Global Ecology and Biogeography, 22: 834-845.    

  • Mertens, R. (1971) -  Die Herpetofauna Südwest-Afrikas. -  Abhandlungen der Senckenbergischen Naturforschenden Gesellschaft, 529: 1-110.  

  • Monard A. (1937) -  Contribution à l’Herpétologie d’Angola. -  Arquivos do Museu Bocage, 8: 19-154.  

  • Parker, H.W. (1936) -  Dr. Karl Jordan`s Expedition to South-West Africa and Angola: Herpetological Collections. -  Novitates Zoologicae, London, 40: 115-146.  

  • Pianka, E.R. (1971) -  Lizard Species Density in the Kalahari Desert. -  Ecology, 52 (6): 1024-1029.    

  • Pietersen, D.W. & Pietersen, E.W. & Haacke, W.D. (2013) -  First herpetological appraisal of the Parque Nacional de Banhine, Gaza Province, southern Mozambique. -  Annals of the Ditsong National Museum of Natural History, 3: 153-163.    

  • Schmidt, A. (1997) -  Jugendentwicklung, ökologische Aspekte und Verbreitung der südafrikanischen Eidechse Heliobolus lugubris. -  Zusammenfassung DGHT Jahrestagung Dormagen vom 10.-14.9.1997. S. 10.  

  • Schmidt, A.D. (1995) -  Mimikry zwischen Wüsteneidechsen und Wüstenlaufkäfern.  -  elaphe N.F., Rheinbach, 3 (3): 74-75.  

  • Schmidt, A.D. (1996) -  Untersuchungen zur Biologie, Ökologie und Verbreitung von südafrikanischen Eidechsen (Reptilia: Lacertidae: Heliobolus lugubris) und Laufkäfer (Coleoptera: Carabidae: Anthia/Thermophilium) als grundlage für eine Mimikry-Hypothese. -  Diplomarbeit Universität Bonn. 168 S.  

  • Schmidt, A.D. (1997) -  Mimikry zwischen Wüsteneidechsen und Wüstenlaufkäfern – eine Überlebensstrategie in extremer Umwelt. -  Reptilia, Münster, 2 (4): 22-24.  

  • Schmidt, A.D. (1997) -  Zur Verbreitung, Biologie und Ökologie von Heliobolus lugubris A. SMITH, 1838 (Sauria, Lacertidae) im südlichen Afrika. -  Salamandra, Rheinbach, 33 (4): 269-288.    

  • Schmidt, A.D. (1997) -  Phänotypische, ethologische und ökologische Unterschiede zwischen juvenilen und adulten Heliobolus lugubris A. SMITH 1838 und deren biologischer Hintergrund. -  Senckenbergiana biologica, Frankfurt/Main, 77 (1): 1-13.  

  • Schmidt, A.D. (1997) -  Zur wachstumsbedingten Entwicklung von Färbung und Biometrie bei Heliobolus lugubris (A. SMITH, 1838) (Sauria, Lacertidae). -  Sauria, Berlin, 19 (2): 35-45.  

  • Schmidt, A.D. (2004) -  Mimikry zwischen Eidechsen und Laufkäfern. -  Frankfurt/Main (Frankfurter Beiträge zur Naturkunde Bd. 23, Edition Chimaira). 374 S.  

  • Schneider, M.F. & Buramuge, V.A. & Aliasse, L. & Serfontein, F. (2005) -  Checklist of Vertebrates of Mozambique. -  Universidade Eduardo Mondlane Faculdade de Agronomia e Engenharia Florestal Departamento de Engenharia Florestal Maputo, Moçambique. 227 pp.  

  • Sclater, W.L. (1898) -  List of the Reptiles and Batrachians of South Africa, with descriptions of New Species. -  Annals of the South African Museum, Cape Town, South Africa, 1 : 95-111.  

  • Seligmann, H. (2000) -  Evolution and ecology of developmental processes and of the resulting morphology: directional asymmetry in hindlimbs of Agamidae and Lacertidae (Reptilia: Lacertilia). -  Biological Journal of the Linnean Society, 69 (4): 461–481.    

  • Smith, A. (1838) -  Contributions to the Natural History of Southern Africa. Art. VIII. -  Magazine of natural history, London, 2 (14): 92-94.  

  • Smith, A. (1849) -  Tropidosaura dumerilii -  Illustrations of the zoology of South Africa, Ill, Appendix: 7.  

  • Sternfeld, R. (1911) -  Die Fauna der deutschen Kolonien. -  Zoologisches Museum Berlin, Reihe IV: Deutsch-Südwestafrika, Heft 2: Die Reptilien (außer den Schlangen) und Amphibien von Deutsch-Südwestafrika.  

  • Szczerbak, N.N. (1975) -  Catalog of the African Sand lizards. -  Naukova Dumka, National Academy of Sciences of Ukraine, Kiev. 83 pp.  

  • Szczerbak, N.N. (1989) -  Catalogue of the African Sand Lizards (Reptilia: Sauria: Eremiainae: Lampreremias, Pseuderemias, Taenieremias, Mesalina, Meroles). -  Herpetozoa, Wien, 1 (3/4): 119-132.    

  • Tornier, G. (1900) -  Neue Liste der Crocodilen, Schildkröten und Eidechsen Deutsch-Ost-Afrikas. -  Zoologische Jahrbücher. Abteilung für Systematik, Geographie und Biologie der Tiere, Jena, 13: 579-681.  

  • Turner, A.A. (2014) -  Heliobolus lugubris (A. Smith, 1838) Bushveld Lizard. -  In: Bates, F.M., Branch, W.R., Bauer, A.M., Burger, M., Marais, J., Alexander, G.J. & M.S. de Villiers (Eds): Atlas and Red List of the Reptiles of South Africa, Lesotho and Swaziland. - Sanbi, Pretoria. p. 160.   

  • Verwaijen, D. & Van Damme, R. (2007) -  Does foraging mode mould morphology in lacertid lizards? -  Journal of Evolutionary Biology, 20 (5): 1950-1961.    

  • Werner, F. (1908) -  Reptilien und Amphibien.  -  Sammlung Göschen, Das Tierreich Band III, Leipzig, 180 pp.  

  • Werner, F. (1910) -  Zoologische und Anrtopologische Ergebnisse einer Forschungsreise im Westlichen und Zentralen Südafrika, ausgeführt in den Jahren 1903-1905. Vierter Band. Systematik und Tiergeographie. Reptilia et Amphibia. Familia Lacertidae. -  Denkschriften der Medichinisch-Naturwissenschaftlichen Gesellschaft zu Jena, 16: 329-340.  

  • Whiting, M.J. (1996) -  Lacertidae: Heliobolus lugubris, bushveld lizard, copulation. -  African Herp News, Bloemfontein, South Africa, 25: 39  

  • Wyk, J.H. van (1984) -  Die Reptiele van die Oranje-Vrystaat. -  Culna, 27: 10-15.